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IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

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7924 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
OLD 1955 BK II 293 101 10 - BK Man 1955 104 - Unknown II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 20. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Perissodactyla Hippomorpha Equidae Equinae Stylohipparion sp. No
G. 356 102 5 - FLK MAN, L/R-P, L/R-M, L/R-C, L/R-I 1960 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 23. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Promesochoerus mukiri Holotype No
FLK NN I 701 103 6 - FLK NN C 104 - Unknown I middle 2 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 24 next to modern Phacochoerus specimens. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Promesochoerus mukiri Paratype No
6084 104 7 - FLK North C 104 - Unknown I upper 1 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Two upper canines presumed to be female. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Promesochoerus mukiri Paratype No
FLK N I 1235 105 7 - FLK North CRA, MAX, P, M 1961 104 - Unknown I upper 5 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 25. It is the same individual as FLK N I 1236. The specimen is crushed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Ectopotamochoerus dubius Holotype No
FLK N I 1236 106 7 - FLK North MAN, P, M 1961 104 - Unknown I upper 6 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. FLK N I 1235 and 1236 are two parts of the same individual and are both considered the holotype. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Ectopotamochoerus dubius Holotype No
FLK NN I 177/1960 107 6 - FLK NN CRA, MAX, P, M, C 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle 3 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 28. This skull is "very crushed."Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Potamochoerus intermedius Holotype No
FLK N I 335 108 7 - FLK North R-MAN , R-P , R-M , R-C 1960 104 - Unknown I upper 1, 2, 3 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Specimen consists of a right mandible that is missing incisors. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Suiformes Suoidea Suidae Tetraconodontinae Notochoerus euilus Holotype. At the time of publication, it was considered along with FLK N I 441 to be the holotype of N. eulius. No
FLK N I 441 109 7 - FLK North MAN 1960 104 - Unknown I upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Suiformes Suoidea Suidae Tetraconodontinae Notochoerus euilus Holotype - The author does not give distinction between this specimen and FLK N I 335. Both seem to be mandibles, perhaps each side of the same individual. At the time of publication, it was considered along with FLK N I 335 to be the holotype of N. eulius. No
OVPP-Tapinochoerus 1 110 5 - FLK R-MAN, R-P, R-M 1959 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 30. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Phacochoerini Tapinochoerus sp. No
OLD 159 111 5 - FLK M 1959 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 31. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Phacochoerini Tapinochoerus No
OVPP-Hippopotamus 1 112 CRA, MAX, MAN 104 - Unknown I, II, IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The author mentions several specimens from beds 1, 2, and 4. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Hippopotamoidea Hippopotamidae Hippopotaminae Hippopotamus gorgops No
FLK N I 637 114 5 - FLK HOR 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle upper layers in the middle level. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Giraffoidea Giraffidae Palaeotraginae Okapia stillei Gentry and Gentry (1978) later identified this specimen as Parmularius altidens. No
OVPP-Giraffa 1 115 11 - HWK SKEL 104 - Unknown II lower Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This is a complete skeleton. Some parts are pictured in plates 33, 34, and 35. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Giraffoidea Giraffidae Giraffinae Giraffini Giraffa gracilis Syntype No
M. 21466 116 MAN, P, M 106 - Natural History Museum UK (?) II, IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 36 and 37. The author indicates this species is well represented in Beds 2 and 4, suggesting other materials may be available in the BMNH. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Giraffoidea Giraffidae Giraffinae Giraffini Giraffa jumae Holotype No
M. 21470 117 12 - CK R-HOR-f 1935 106 - Natural History Museum UK (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros sp. This specimen was reclassified as Tragelaphus stepsiceros by Gentry and Gentry (1978). No
M. 21461 118 13 - RK CRA, R/L-HOR, R-I, R-P, R-M 1931 106 - Natural History Museum UK (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 38 and 39. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros grandis Holotype. Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros grandis. No
M. 14549 119 HOR-f 1932 106 - Natural History Museum UK (?) I, II, III Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Leakey (1965) indicated this specimen was found on the surface of Bed III, though the museum's label indicates Bed I and markings on the horn core indicate Bed II. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros grandis Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros grandis. No
M. 21469 120 12 - CK R-HOR-f 1935 106 - Natural History Museum UK (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. No information is given about elements preserved. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros grandis Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros grandis. No
OVPP-Tragelaphus 1 121 14 - HWK E CRA, HOR 1959 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II lower 2 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 40, 41. The cranium and horn cones were found near the base of Bed 2. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros maryanus Holotype - Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros maryanus. No
OVPP-Tragelaphus 2 122 6 - FLK NN CRA, HOR 1960 104 - Unknown I middle 1, 2, 3 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Other specimens are present (Leakey, 1971) but are not described. The cranium and horn cones were found near the base of Bed 2. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros maryanus Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros maryanus. No
068/5813 123 58 - KK CRA 1959 104 - Unknown II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The author indicates that specimens are common in Bed I, lower Bed II, and early in middle Bed II (Gentry and Gentry, 1978, a). The specimen # is provided by Gentry and Gentry (1978). The cranium and horn cones were found near the base of Bed 2. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros maryanus Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros maryanus. No
F. 3665 P.P.T. 4 124 16 - LK CRA, L-HOR-f, R-HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Face and maxillae are missing. The holotype is pictured in plates 43 and 44. It was found on the surface between RK and LK. This eland is poorly represented at Olduvai. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Taurotragini Taurotragus arkelli Holotype No
M. 14947 125 17 - PLK FRLT-f, L/R-HOR-f 106 - Natural History Museum UK (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Specimen is pictured in plate 46. According to the author, it is "in bad shape." Gentry and Gentry suggest this specimen probably came from the top of Bed 2 at PLK. 130 - L.S.B. Leakey 1936 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Bularchus arok Holotype - Gentry and Gentry (1978) reclassidfied this specimen as Pelorovis oldowayensis. No
M. 14948 126 CRA-f, L-HOR-f 106 - Natural History Museum UK (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Bularchus arok Paratype - Gentry and Gentry (1978a) renamed this specimen as P. oldowayensis. No
OVPP-Bularchus 1 127 10 - BK SKULL, MAX, CRA, HOR 104 - Unknown II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Several specimens including crania and post-cranial elements have been found in BK II. One example is pictured in plate 48. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Bularchus arok Gentry and Gentry (1978a) renamed this specimen as P. oldowayensis. No
M. 21451 128 18 - VEK CRA-f, R-HOR-f 1935 106 - Natural History Museum UK (?) III, IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This holotype was found at the junction of Beds 3 and 4. It is pictured in plates 49 and 50. [This species is documented at SHK II, FLK N I, and BK II (Leakey, 1971).] 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Gorgon olduvaiensis Holotype - Gentry and Gentry (1978) reclassified this specimen as Connochaetes taurinus olduvaiensis. No
BK II 1955 P.P.F. 1 129 10 - BK R-FRO, R-HOR 1955 106 - Natural History Museum UK (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen was incorrectly stated by Leakey (1965) that its museum number is M. 21452 but this was corrected by Gentry and Gentry (1978a). It is pictured in Leakey (1965) plate 52 (lower) and in Gentry and Gentry (1978a) plate 16, figure 3. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Gorgon olduvaiensis This specimen was later reclassified as Connochaetes taurinus olduvaiensis by Gentry and Gentry (1978). No
OVPP-Gorgon 1 130 L-FRO, L-HOR-f 106 - Natural History Museum UK (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is in the BMNH collections but a specimen number is not given. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Gorgon olduvaiensis No
OVPP-Gorgon 2 131 10 - BK HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Gorgon olduvaiensis No
P.P.R. 1 132 10 - BK FRLT, L/R-HOR 1953 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Reduncinae Reduncini Kobus idet. This specimen was reclassified by Gentry and Gentry (1978) as Kobus kob. No
579/1957 133 19 - SHK FRO-f, PAR-f, R-HOR 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Reduncinae Reduncini Kobus sp. No
FLK N I 1039 134 7 - FLK North FRLT, L/R-HOR-f 104 - Unknown I upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition, 273 - Reck expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Reduncinae Reduncini Indet. This specimen was reclassified by Gentry and Gentry (1978) as Antidorcas recki. No
M. 14561 135 20 - JK 1 HOR 1936 106 - Natural History Museum UK (?) III Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 293 - A.T. Hopwood 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Damaliscus niro Holotype - This specimen was reclassified by Gentry and Gentry (1978) as Damaliscus niro. No
OVPP-Hippotragus 1 136 HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Several specimens are present in the Nairobi Museum's collections. Three are pictured in plate 55. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus niro No
53/282 137 FRO, L/R-HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus sp. No
F. 3662 068/5812 P.P.T.2 138 CRA-f, L-HOR, R-HOR-f 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 56 and 58. The adhering matrix suggests this specimen must come from above the Lemuta member. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas Holotype No
1957.56 P.P.T.3 139 18 - VEK CRA-f, L/R-HOR 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper Upper Villafranchian Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 59 and 60. It is from a female. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas Paratype No
M. 21448 140 FRO, HOR 1931 106 - Natural History Museum UK (?) II surface Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 61. 293 - A.T. Hopwood 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas No
F. 992 141 HOR-f 1941 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas No