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IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

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410 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
OLD 15740 10156 6 - FLK NN PMAX I Layer 3 Brodkorb P., Mourer-Chauviré, C. 1984. Fossil owls from early man sites of Olduvai Gorge, Tanzania. Ostrich; Journal of African Ornithology 55:17-27. PARATYPE. "Paratypes: The following specimens are from layer 3 of locality FLK NNI and can be attributed to the same individual as the holotype: 15740, premaxillae; 3301, lower part of right coracoid. 15364 and 15473, proximal and distal parts of left femur; 16377, distal part of right femur; 15701, distal part of left tibiotarsus; 15776, left fibula; 16262, proximal part of right tarsometatarsus; 15730, distal part of left tarsometatarsus; 15942, 15946, 16050, 16171, 16174, 16176 to 16180, pedal phalanges. Place of conservation of the material: The fossils are currently on deposit at the University of Florida, Department of Zoology, and will later be returned to Tanzania" (Brodkorb and Mourer-Chauvire, 1984:17, 19). Image notes: (Scale = 2 cm) A-B: Left coracoid, holotype, OLD 15802, FLK NNI, layer 3, A anterior face, B posterior face. C: Right tarsometatarsus, proximal part, paratype, OLD 16262, FLK NNI, scree, anterior face. D: Left tarsometatarsus, distal part, paratype, OLD 15730, FLK NNI, layer 3, anterior face. E-F: Left femur, proximal half, paratype, OLD 15364, FLK NNI, layer 2+3, E anterior face, F POSterior face. G-H: Left femur, distal part, paratype, OLD 15473, FLK NNI, layer 3, G anterior face, H posterior face. I: Left tarsometatarsus, almost complete, OLD 11574, FLK NI, layer 1, anterior face. J-K: Left femur, lacking the distal part, OLD 11586, FLK NI, layer 2, J anterior face, K posterior face. Unknown Aves Strigiformes Strigidae Bubo leakeyae No
OVPP-Bitis 8 1935 MAX-f 104 - Unknown I Rage J. 1973. Fossil snakes from Olduvai, Tanzania. In: Fossil vertebrates of Africa, vol. 3:1-6. 149 - J. C. Rage Unknown Reptilia Squamata Viperidae Bitis No
OVPP-Bitis 9 1936 MAX-f 104 - Unknown I Rage J. 1973. Fossil snakes from Olduvai, Tanzania. In: Fossil vertebrates of Africa, vol. 3:1-6. 149 - J. C. Rage Unknown Reptilia Squamata Viperidae Bitis No
OVPP-Bitis 13 1940 5 - FLK L-MAX 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Rage J. 1973. Fossil snakes from Olduvai, Tanzania. In: Fossil vertebrates of Africa, vol. 3:1-6. This specimen is pictured in Plate 1, figures 5-8. 149 - J. C. Rage Yes Reptilia Squamata Viperidae Bitis olduvaiensis Holotype No
OVPP-Bitis 14 1941 5 - FLK MAX-f 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Rage J. 1973. Fossil snakes from Olduvai, Tanzania. In: Fossil vertebrates of Africa, vol. 3:1-6. This specimen is pictured in plate 1, figure 9. 149 - J. C. Rage Unknown Reptilia Squamata Viperidae Bitis olduvaiensis No
OVPP-Muridae 1 35 5 - FLK P, M, I, C, MAN, MAX 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Several unidentified murids 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae No
OVPP-Heterocephalus 2 3008 7 - FLK North MAX-f, M, (N=7) 104 - Unknown I upper Denys, C. 1989. A new species of bathyergid rodent from Olduvai Bed I (Tanzania, Lower Pleistocene). Neue Jahrbuecher Geologica Palaeontologica 5:257-264. The author mentions 7 maxillae with associated molars. 147 - C. Denys Yes Mammalia Rodentia Bathyergidae Heterocephalinae Heterocephalus jaegeri No
01. R. L3-95 3121 6 - FLK NN R-MAX-f, M 104 - Unknown I 3 Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 1, figure A. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Oenomys olduvaiensis Holotype No
01. R. L3-88 3122 6 - FLK NN R-MAX-f, M 104 - Unknown I 3 Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 1, figure A. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Grammomys sp. No
01. R. K-199 3124 5 - FLK R-MAX-f, M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figures A and B. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus Holotype No
01. R. K-533 3125 5 - FLK R-MAX-f, M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figure E. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Aethomys lavocati Holotype No
01. R. M5-222 3126 7 - FLK North R-MAX-f, M 104 - Unknown I 5 Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figure C. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Pelomys dietrichi Holotype No
01. R. L2-2 3127 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 3, figures A and B. This species is found at FLK NI, layers 1, 2, 3, 4, 5, and 6; FLK NNI, layers 1, 2, and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Mus petteri Holotype No
01. R. L5-58 3128 7 - FLK North L-MAX-f, M 104 - Unknown I 5 Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 3, figures G and H. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. K-120 3129 5 - FLK L-MAX-f, M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 1, figure D. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-314 3130 5 - FLK R-MAX-f, M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 3, figure D. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Zelotomys leakeyi Holotype No
01. R. K-20 3150 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M1-29 3159 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M4-35 3165 R-MAX-f, PAL, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M4-36 3166 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M4-37 3167 R-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M4-38 3168 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M5-58 3188 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M5-60 3190 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M5-61 3191 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M5-62 3192 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M5-63 3193 R-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M5-64 3194 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. M4-541 3218 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NI, layers 2, 3, 4, 5, and 6; FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. K-90 3219 R-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Oenomys olduvaiensis No
01. R. K-90 3220 R-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Oenomys olduvaiensis No
01. R. L3-97 3226 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Oenomys olduvaiensis No
01. R. L3-98 3227 R-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Oenomys olduvaiensis No
01. R. L3-99 3228 R-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK NNI, layers 2 and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Oenomys olduvaiensis No
01. R. K-116 3245 L-MAX, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-116 3246 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-117 3247 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-118 3248 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-119 3249 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-122 3251 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 1, figure F. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No