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IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

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7924 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
01. R. K-171 3300 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-172 3301 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-173 3302 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-174 3303 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-175 3304 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-176 3305 L-M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-177 3306 L-M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-178 3307 L-M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-179 3308 R-M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-180 3309 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-181 3310 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-182 3311 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. L2-183 3312 R-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. L2-184 3313 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. L2-185 3314 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. L2-186 3315 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. L3-187 3316 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M2-188 3317 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M2-189 3318 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M2-190 3319 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M2-191 3320 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M3-192 3321 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M3-193 3322 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M4-194 3323 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M4-195 3324 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M4-196 3325 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M4-197 3326 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M4-198 3327 L-M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. L3-538 3328 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. M1-539 3329 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-540 3330 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layers 1, 2, 3, and 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Thallomys quadrilobatus No
01. R. K-200 3331 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figure B. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. K-201 3332 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. K-202 3333 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. K-203 3334 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. K-204 3335 L-M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. L2-205 3336 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. L3-206 3337 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. L3-207 3338 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
01. R. L3-208 3339 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No