Search the database

IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

Search Results

7924 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
OH 63 3778 6 - FLK NN M 1988 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I "Found near the OH 7 site" -R. Clarke 2011. Specimen is a lower molar. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus No
OH 64 3779 81 - HWK W TOOTH 1989 104 - Unknown II Lowermost 20 mm above Tuff IF in "basal waxy clay". Clarke, R.J. Published online 4 May 2012. A Homo habilis maxilla and other newly-discovered hominid fossils from Olduvai Gorge, Tanzania. Olduvai Special issue, JHE Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 65 3780 82 - 64 MAX , TOOTH 1995 127 - National Natural History Museum, Arusha, Tanzania I Middle/Upper Blumenschine, R. J., Peters, C. R., Masao, F. T., et al. 2003. Late Pliocene Homo and Hominid Land Use from Western Olduvai Gorge, Tanzania. Science 299, 1217-1221. Author describes specimen as "complete maxilla with all teeth and lower face of adult". Specimen is pictured in images 1-11. (Images from R. Clarke 2011). Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Large habilis No
OH 66 3781 7 - FLK North R-P 1998 127 - National Natural History Museum, Arusha, Tanzania I Upper Surface. Tooth is lower P4. Specimen is pictured on the far right in image 1. Specimen is pictured in bottom left in image 2. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 67 3782 14 - HWK E M-f 2001 127 - National Natural History Museum, Arusha, Tanzania II Lowermost Surface. Upper adult fragment. Specimen is pictured on top right in image 1. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homo or Australopithecus No
OH 68 3783 18 - VEK Mand-f, R-P-f 2001 127 - National Natural History Museum, Arusha, Tanzania II Lower Mandible fragment with unerupted P4 and root of M1. Specimen is pictured in the middle in image 1. Specimen is shown in the middle of the top row in image 2. Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus africanus OH 13-like No
OH 69 3784 80 - MCK L-P 2002 127 - National Natural History Museum, Arusha, Tanzania II Lower Lower P4. Specimen is pictured on left in image 1. Specimen is pictured on left in image 2. Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis sunsu lato No
OH 70 3785 80 - MCK L-M-f 2002 127 - National Natural History Museum, Arusha, Tanzania II Lower Distrobuccal portion of upper M1 crown. Specimen is pictured on bottom right of image 1. Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus africanus OH 24-like No
OH 71 3786 24 - DK TOOTH 2003 127 - National Natural History Museum, Arusha, Tanzania I Lower “Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Specimen is shown. Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 72 3787 24 - DK TOOTH 2004 127 - National Natural History Museum, Arusha, Tanzania I Lower “Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 73 3788 81 - HWK W L-P 2006 127 - National Natural History Museum, Arusha, Tanzania I Upper Surface. Clarke, R.J. Published online 4 May 2012. A Homo habilis maxilla and other newly-discovered hominid fossils from Olduvai Gorge, Tanzania. Olduvai Special issue, JHE Left P3. Specimen is pictured third from the left bottom in image 1. Specimen is shown on left in image 2. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis sensu stricto No
OH 74 3789 72 - FLK Maiko Gully L-M 2007 127 - National Natural History Museum, Arusha, Tanzania I ID above Surface. Left M2, unerupted. Specimen is pictured second from the bottom left in image 1. Specimen is shown on right in image 2. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 75 3790 5 - FLK R-HUM 2009 127 - National Natural History Museum, Arusha, Tanzania I Zinjanthropus Surface. OLAPP/Earthwatch”Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) No Mammalia Primates Anthropoidea Haplorhini Hominoidea hominidae A. cf. africanus/H. habilis OH 62-like No
OH 76 3791 55 - FLK W R-M 2010 127 - National Natural History Museum, Arusha, Tanzania II Upper? Surface. “Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Right upper M1 with roots. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 77 3792 14 - HWK E L-M-f 2010 127 - National Natural History Museum, Arusha, Tanzania II Lowermost Surface. OLAPP/Earthwatch”Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Left upper M2 crown. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus africanus No
OH 78 3793 10 - BK H-f 2007 127 - National Natural History Museum, Arusha, Tanzania II Lower-Middle? Surface. OLAPP/Earthwatch”Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Humerus shaft, subaerial weathering. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus africanus OH 75-like No
OH 79 3794 49 - BK E CRA-f 2007 127 - National Natural History Museum, Arusha, Tanzania II Lower-Middle Surface. OLAPP “Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011)/Earthwatch Weathered. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 80 3796 10 - BK R/L-P , L-M , R/L-I , L-HUM-f , R-RAD-f , R-FEM-f , TIB-f 2009 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II 4 Upper Domínguez-Rodrigo M, et al. 2013. First Partial Skeleton of a 1.34-Million-Year-Old Paranthropus boisei from Bed II, Olduvai Gorge, Tanzania. PLoS ONE 8(12): e80347. Distal portion of a left humerus diaphysis. Partial right radius with head, proximal metaphysis and partial diaphysis. Right partial femur, diaphysis in three refitting pieces. Tibia midshaft fragment assigned to OH 80 based on close "spatial association, comparable anatomical ruggedness and similar state of preservation to the other OH 80 postcranial fossils" (Dominiguez-Rodrigo et al., 2013). 292 - Proyecto Olduvai Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Australopithecus boisei Anatomically modern No
OH 1968 3797 40 - JK CRA 1968 125 - Germany Ndutu Lower Surface. Von Zieten RRR. 2009. Preliminary report on a new Olduvai hominid from 1968: an Anatomically-Modern-Man from Upper Ndutu Bed. Retrieved from http://www1.uni-hamburg.de/ Anatomically modern. Complete skull. Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sapiens Anatomically modern No
MLK 068/6514 3803 34 - MLK CRA-f, P, M, (n=3) II Lower From horizon above Lemuta Member of Lowed Bed II Leakey MG and Leakey REF. 1973. New large Pleistocene colobinae (Mammalia, Primates) from East Africa. Fossil Vertebrates of Africa Vol. 3, 121-38. Similar to colobine specimen KNM-ER 398 in cranial dimensions. Maxillary dentition represented by P4, M2, M1, M2. Crania is represented by multiple fragments, and some fragments are missing. Large canine socket of right maxilla indicates specimen is male. Specimen is pictured in Plate 3. 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Either Cercopithecinae or Cynopithencinae. Author states that the cranium is typical of Cercopithecinae but that the dental morphology resembles Cynopithecinae. No
Old. 69 S. 194. rt. M2 3919 III 202/TII Leakey, M. G. 1982. Extinct large colobines from the Plio-Pleistocene of Africa. American Journal of Physical Anthropology 58:153-172. Most complete specimen of this species. 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Colobinae Colobini Cercopithecoides kimeui No
5349 3933 P I/II Petter, G. 1973. Carnivores Pleistocenes du ravin d’Olduvai (Tanzanie). Fossil vertebrates of Africa 3:43-100. Specimen is lower P4. Author lists specimen in an assemblage that is described as coming from FLK N, FLK NN, KK, BK, and DK but does not specifically assign site. 157 - L. Hlusko Unknown Mammalia Primates No
2609 3943 MAX, P, (n=4), M, (M=2) II Petter, G. 1973. Carnivores Pleistocenes du ravin d’Olduvai (Tanzanie). Fossil vertebrates of Africa 3:43-100. Author lists specimen in an assemblage that is described as coming from FLK N, FLK NN, and MNK but does not specifically assign site. Specimen is pictured in image 1. 157 - L. Hlusko 2012/6/28 Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae No
B 44 3952 27 - MNK L-CUB II Petter, G. 1973. Carnivores Pleistocenes du ravin d’Olduvai (Tanzanie). Fossil vertebrates of Africa 3:43-100. 157 - L. Hlusko 2012/6/28 Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Cercopithecinae Papionini Theropithecus No
OLD 1472 3694 63 - JLK MAN, M, P 1957 104 - Unknown IV Leakey, L., and Whitworth, T. 1958. Notes on the genus Simopithecus, with a description of a new species from Olduvai. This specimen is pictured in plate 4, figure 1; plate 5, figure 1; plate 6, figure 1; plate 8, plate 9, and plate 10. All descriptions of this specimen indicate that the pictured specimen is the holotype. In plate 4, however, the description reads female S. jonathani so it is unclear whether this is the same specimen or not. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Cynocephalinae Simopithecus jonathani Holotype No
OLD 580 3695 22 - SC MAN-f, M 1957 104 - Unknown II Leakey, L., and Whitworth, T. 1958. Notes on the genus Simopithecus, with a description of a new species from Olduvai. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Cynocephalinae Simopithecus jonathani No
068/6514 3711 34 - MLK CRA, MAX-f, R/L-P, R/L-M 104 - Unknown II middle Leakey, M. G., and Leakey, R. E. F. 1973. Pleistocene Colobinae from East Africa. Leakey, L. S. B., Savage, R. J. G., Coryndon, S. C., eds. In: Fossil Vertebrates of Africa,vol. 3. London: Academic Pr This specimen is pictured in plate 3. 154 - Leakey and Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Colobinae Colobini Cercopithecoides kimeui Holotype No
OH 5 3713 5 - FLK 283 - 45 CRA , MAX , M , I , C , P 17 July - 6 August 1959 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Middle (Zinj.) 22 feet below the upper limit of Bed I (Tuff IF), approximately 20 feet above basalt. Leakey, L. S. B. 1959. A new fossil skull from Olduvai. Nature 184:491-493. Holotype. Almost complete cranium. Adolescent. One reconstructed fossil found at House of Culture, Dar es Salaam, Tanzania. Specimen previously housed at National Museum of Kenya. Additional pieces from NMK unpublished. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Zinjanthropus boisei Holotype. This specimen was reclassified as Paranthropus boisei by Robinson (1960). No
OH 1 3717 13 - RK 278 - 9 SKEL 1913 116 - Bayerische Staatssammlung für Paläontologie und Geologie (BSPG), Munich, Germany Ndutu Intrusive burial Hans-Reck. 1913, Oldoway-Expedition Monograph; LSB Leakey 18 March 1933 Partial skeleton. Intrusive burial post-dating primary Olduvai sequence. 144 - H. Reck 273 - Reck expedition Unknown Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sapiens No
OH 2 3718 27 - MNK 279 - 88 CRA-f 1935 106 - Natural History Museum UK (?) III/IV Lowest horizon of Bed IVa. Surface. Found at head of MNK Gully where Beds II and IV are exposed. Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Two cranial vault fragments. Specimen previously housed at National Museum of Kenya. Not found at House of Culture, Dar es Salaam, Tanzania. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus Similar to OH 9 and OH 12. No
OH 4 3720 8 - MK 281 - 11 P , M , MAN 1959 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Lower Surface. Leakey, L.S.B., Tobias, P.V. & Napier, J.R. 1964. A new species of the genus Homo from Olduvai Gorge. Nature 202(4927): 7-9. Paratype. Two isolated teeth (lower molar and upper premolar) and one M3 in a mandible. Found in small block of consolidated tuff. Specimen previously housed at National Museum of Kenya. 282 - H. Mukiri 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype No
OH 3 3719 10 - BK 280 - 94 d-C , L-d-M 1955 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Upper Upper part of BK channel which cuts into Tuff IID Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. In the first publication of these specimens (Leakey, LSB, 1958), Leakey described the teeth as a deciduous lower canine and a left deciduous upper M2. "This view was not shared by Robinson, von Koenigswald and others, who preferred to regard it as either a permanent or a deciduous upper molar of an australopithecine character" (Leakey, MD, 1970:226). Specimen previously housed at National Museum of Kenya. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus Referred to as species boisei in Tobias (Olduvai Gorge volume IV, 1991). No
OH 6 3721 5 - FLK 283 - 45 CRA-f , M-f , L-P , L-TIB-f , L-FIB-f , 1959/60 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Middle (Zinj.) Surface. Cranial and dental fragments found in surface soil from slope immediately below Zinj. site; assumed to be from Zinj. level (Leakey, MD, 1971:227). TIB-f and FIB-f from Zinj. level. Davis, P. R. 1964. Hominid fossils from Bed I, Olduvai Gorge, Tanganyika. A tibia and fibula. Nature. 201, 967. From Tobias (1991): OH 35 tibia and fibula attributed to same individual as OH 6 but appear to be more mature. It is unclear if OH 35 tibia and fibula are the same tibia and fibula originally listed under OH 6. Leakey, MD, 1971 describes CRA and dental fragments as having been found in surface soil, but Leakey, Tobias, and Napier, 1964 state "some [were] found in situ [and] some on the surface" (:9). Unworn lower L-P3. Partially developed roots of upper M1 or 2. Six cranial vault fragments. Tibia and fibula fragments first assumed to be associated with Zinjanthropus OH5, but should be considered associates with OH 6 following morphological study (Leakey, MD, 1971:227). and skull fragments. Juvenile. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype. TIB-f and FIB-f not given taxonomic designation in original publication (Davis, 1964) but named as a paratype in Leakey, Tobias, and Napier, 1964. No
OH 7 3722 6 - FLK NN 283 - 45 MAN , L-PAR , R-PAR-f , M , OCC-f , L/R-TEM-f , PHX , CAR 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 On surface of clay immediately overlying Tuff IB. Leakey, L.S.B., Tobias, P.V. & Napier, J.R. 1964. A new species of the genus Homo from Olduvai Gorge. Nature 202(4927): 7-9. Holotype. "Jonny's Child". Juvenile. Mandible with partial dentition and unerupted R/L-M3, almost complete left parietal and fragments of right parietal, left and right petrous portions (temporal) , "sundry small skull fragments ... [and] twenty-one bones of the hand, including the trapezium, scaphoid and a number of phalanges" (Leakey, MD, 1971:228). "One upper molar, a terminal phalanx, and a broken capitate bone, found on the surface, probably also belong to this individual" (:226). Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Holotype No
OH 8 3723 6 - FLK NN 283 - 45 PHX , CAR , TAR , CLA , AST , CAL , CUNE , CUB , NAV , M-f 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. The molar-f was found on the surface and "probably also belongs to OH 8" (Leakey MD, 1971:228). Twelve associated foot bones including astragalus (has carnivore tooth marks (Leakey MD, 1971), calcaneus, cuboid and navicular, cuneiforms 1- 3, proximal ends with shafts, five metatarsals. Proximal ends of one metacarpal and three finger phalanges. One radius shaft. Clavicle with damaged articular ends. Foot articulates "almost perfectly with the OH 35 leg" (Susman, 1983:372). 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype No
OH 9 3724 71 - LLK 284 - 46 CALV 2 December 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Upper II D Surface Leakey LSB. 1966. Homo habilis, Homo erectus and the Australopithecines. Nature, 209(5030), 1279–1281. Calvaria. "Chellean Man." 264 - Leakey family expedition No Yes. Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 10 3725 7 - FLK North 283 - 45 R-PHX 1961 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper (level 5) 3.5 feet below marker Tuff IF at top of Bed I Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Terminal first phalanx. Specimen previously housed at National Museum of Kenya. 154 - Leakey and Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini No
OH 11 3726 24 - DK 285 - 13 PAL , L-MAX-f , TOOTH-f 1962 122 - National Museum and House of Culture, Dar es Salaam, Tanzania Ndutu Lower Found in stream bed at bottom of gully near DK excavations. See curatorial notes for details. Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left half of palate and maxillary arch, crowns of teeth not preserved. "It is thus impossible to assign to any stratigraphic level, although the condition of the bone, which bears a pinkish tinge, is similar to that seen in fossil bones from a conglomerate at the head of the DK gully belonging to the lower Ndutu Beds" (Leakey MD, 1971:230). Specimen previously housed at National Museum of Kenya. 154 - Leakey and Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sp. No
OH 12 3727 18 - VEK 286 - 86 OCC , L/R-PAR , L/R-TEM-f , CRA-f , L-PAL , L-MAX , TOOTH-f 1962 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV? a Surface. Found on eroded slope of Bed III. L-TEM-f from just below marker Tuff IVa. Leakey, L., and Leakey, M. 1964. Recent discoveries of fossil hominids in tanganyika: at Olduvai and near Lake Natron. Nature 202:5-7. Crowns of teeth not preserved. Bed IVA determined on basis of matrix adhering to L-TEM-f and PAL. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No L-TEM-f & PAL found with matrix adhering. Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 13 3728 27 - MNK 287 - 88 "Skull Site" MAN , MAX , CRA-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle IIB 5 feet above chert horizon Paratpye. "This consists of a sub-adult skull, probably female, with the third molars coming into wear. It is represented by the greater part of the mandible, including the complete dentition, although the crowns of the incisors are damaged and the left ascending ram us is not preserved; the greater part of the maxillae, represented by two fragments both lacking the anterior portions; the greater part of the occipital; the right parietal and temporal; parts of the left parietal and temporal; together with a few frontal fragments and approximately fifty additional pieces which it has not been possible to fit together" (Leakey MD, 1971:231). Specimen previously housed at National Museum of Kenya. 5 pieces found at House of Culture, Dar es Salaam, Tanzania. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype No