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IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

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85 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
OH 1968 3797 40 - JK CRA 1968 125 - Germany Ndutu Lower Surface. Von Zieten RRR. 2009. Preliminary report on a new Olduvai hominid from 1968: an Anatomically-Modern-Man from Upper Ndutu Bed. Retrieved from http://www1.uni-hamburg.de/ Anatomically modern. Complete skull. Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sapiens Anatomically modern No
OH 79 3794 49 - BK E CRA-f 2007 127 - National Natural History Museum, Arusha, Tanzania II Lower-Middle Surface. OLAPP “Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011)/Earthwatch Weathered. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 75 3790 5 - FLK R-HUM 2009 127 - National Natural History Museum, Arusha, Tanzania I Zinjanthropus Surface. OLAPP/Earthwatch”Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) No Mammalia Primates Anthropoidea Haplorhini Hominoidea hominidae A. cf. africanus/H. habilis OH 62-like No
OH 29 3744 76 - JK W M-f 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania III/IVa Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 28 3743 38 - WK 290 - 36 L-PVG-f , L-FEM-f 1970 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV a Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left femoral shaft and left innominate. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 27 3742 37 - HWK EE R-M 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 meters below Tuff IF Surface. Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Lower M3, unworn. "It is not possible to indicate more than broadly the original position of this tooth, since the clays above and below Tuff IF, as well as the Sandy Conglomerate, are all fossiliferous in this areaand yield mammalian teeth in a similar state of preservation to the hominid molar...the tooth was derived from either Upper Bed I or Lower Bed II" (Tobias, 1991:849). Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 72 3787 24 - DK TOOTH 2004 127 - National Natural History Museum, Arusha, Tanzania I Lower “Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 73 3788 81 - HWK W L-P 2006 127 - National Natural History Museum, Arusha, Tanzania I Upper Surface. Clarke, R.J. Published online 4 May 2012. A Homo habilis maxilla and other newly-discovered hominid fossils from Olduvai Gorge, Tanzania. Olduvai Special issue, JHE Left P3. Specimen is pictured third from the left bottom in image 1. Specimen is shown on left in image 2. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis sensu stricto No
OH 74 3789 72 - FLK Maiko Gully L-M 2007 127 - National Natural History Museum, Arusha, Tanzania I ID above Surface. Left M2, unerupted. Specimen is pictured second from the bottom left in image 1. Specimen is shown on right in image 2. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 18 3733 55 - FLK W 283 - 45 PHX 1969 104 - Unknown II Middle- 10 feet above base of Bed II, below Tuff IIC Surface Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Phalanx of the hand. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 22 3737 73 - VEK/MNK R-MAN-f , R-P , R-M , R-I-f , R-C-f 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania III/IV Surface, indeterminate gravel Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Right half of mandible with roots of I1-C and complete P3-M2. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo cf. erectus No
OH 76 3791 55 - FLK W R-M 2010 127 - National Natural History Museum, Arusha, Tanzania II Upper? Surface. “Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Right upper M1 with roots. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 77 3792 14 - HWK E L-M-f 2010 127 - National Natural History Museum, Arusha, Tanzania II Lowermost Surface. OLAPP/Earthwatch”Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Left upper M2 crown. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus africanus No
OH 78 3793 10 - BK H-f 2007 127 - National Natural History Museum, Arusha, Tanzania II Lower-Middle? Surface. OLAPP/Earthwatch”Discovered by an Earthwatch project led by F.T. Masao in 2003 and 2004, respectively, and will be described elsewhere.” -R. Clarke (2011) Humerus shaft, subaerial weathering. No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus africanus OH 75-like No
OH 8 3723 6 - FLK NN 283 - 45 PHX , CAR , TAR , CLA , AST , CAL , CUNE , CUB , NAV , M-f 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. The molar-f was found on the surface and "probably also belongs to OH 8" (Leakey MD, 1971:228). Twelve associated foot bones including astragalus (has carnivore tooth marks (Leakey MD, 1971), calcaneus, cuboid and navicular, cuneiforms 1- 3, proximal ends with shafts, five metatarsals. Proximal ends of one metacarpal and three finger phalanges. One radius shaft. Clavicle with damaged articular ends. Foot articulates "almost perfectly with the OH 35 leg" (Susman, 1983:372). 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype No
OH 13 3728 27 - MNK 287 - 88 "Skull Site" MAN , MAX , CRA-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle IIB 5 feet above chert horizon Paratpye. "This consists of a sub-adult skull, probably female, with the third molars coming into wear. It is represented by the greater part of the mandible, including the complete dentition, although the crowns of the incisors are damaged and the left ascending ram us is not preserved; the greater part of the maxillae, represented by two fragments both lacking the anterior portions; the greater part of the occipital; the right parietal and temporal; parts of the left parietal and temporal; together with a few frontal fragments and approximately fifty additional pieces which it has not been possible to fit together" (Leakey MD, 1971:231). Specimen previously housed at National Museum of Kenya. 5 pieces found at House of Culture, Dar es Salaam, Tanzania. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype No
OH 14 3729 27 - MNK 287 - 88 "Skull Site" CRA-f , FRO-f , R-PAR-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle B Surface. Inferred as same horizon as OH 13 (Leakey MD, 1971). Leakey, L.S.B., Tobias, P.V. & Napier, J.R. 1964. A new species of the genus Homo from Olduvai Gorge. Nature 202(4927): 7-9. Juvenile (sutures unfused). Thin skull. Specimen previously housed at National Museum of Kenya. 11 pieces found at House of Culture, Dar es Salaam, Tanzania. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Juvenile, parietal is thin. No
OH 3 3719 10 - BK 280 - 94 d-C , L-d-M 1955 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Upper Upper part of BK channel which cuts into Tuff IID Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. In the first publication of these specimens (Leakey, LSB, 1958), Leakey described the teeth as a deciduous lower canine and a left deciduous upper M2. "This view was not shared by Robinson, von Koenigswald and others, who preferred to regard it as either a permanent or a deciduous upper molar of an australopithecine character" (Leakey, MD, 1970:226). Specimen previously housed at National Museum of Kenya. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus Referred to as species boisei in Tobias (Olduvai Gorge volume IV, 1991). No
OH 43 3758 6 - FLK NN L-MTT, L-MTT 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 45 3760 6 - FLK NN L-M 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper M1 germ. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 2 3718 27 - MNK 279 - 88 CRA-f 1935 106 - Natural History Museum UK (?) III/IV Lowest horizon of Bed IVa. Surface. Found at head of MNK Gully where Beds II and IV are exposed. Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Two cranial vault fragments. Specimen previously housed at National Museum of Kenya. Not found at House of Culture, Dar es Salaam, Tanzania. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus Similar to OH 9 and OH 12. No
OH 17 3732 72 - FLK Maiko Gully d-M 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Surface Stream bed of Maiko Gully Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 19 3734 29 - FC W 288 - 89 M-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Middle 15 feet above chert horizon Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Half of a very worn molar. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 20 3735 11 - HWK 289 - 44 L-FEM-f 1959 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II or I Lowermost if Bed II, upper if Bed I Surface Day, MH. 1969. Femoral Fragment of a Robust Australopithecine from Olduvai Gorge, Tanzania. Nature. 22: 230-233. Proximal end of large left femur missing head, greater and lesser trochanters damaged. Either Lower Bed II or Upper Bed I (Leakey MD, 1971:233). Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus cf. boisei No
OH 21 3736 7 - FLK North L-M 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper Surface Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Upper left M1, small and unworn. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 23 3738 5 - FLK L-MAN-f , L-P , L-M , 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV/Masek Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left mandible fragment with P4-M2. Specimen previously housed at National Museum of Kenya. Bed IV in Leakey (1971), referred to as having derived from Masek Beds in other publications (e.g., Millard, 2009). 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 24 3739 74 - DK E CRA , M , P 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Lower Surface Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Cranium is crushed. Teeth not specified. From same horizon as OH 4. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 25 3740 268 - 54 L-PAR-f 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV a Surface Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left parietal fragment ca. asterion. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Juvenile. No
OH 26 3741 55 - FLK W R-M 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I/II See curatorial notes. Upper Surface, .6 m below Tuff If at top of Bed I. Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Large unworn right L. M3. "Direct evidence as to the original position of the tooth was ... not obtained but, on the basis of the adhering matrix, which is indistinguishable from the limestone nodules overlying Tuff IF, it can be assumed to have come from the base of Bed II..." (Tobias, 1991:848). Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Found embedded in small nodule of calcareous concretion. Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 36 3751 22 - SC ULNA 1970 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Upper Tuff IID Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Almost complete. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 37 3752 5 - FLK L-MAN , M , M 1971 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Middle Surface, probably from sandy conglomerate in Middle Bed II. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Left half mandible with M1-M2. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 38 3753 22 - SC R-M , I , I 1971 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Upper >IID Fluvo-lacustrine Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Molar is right M2. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 39 3754 37 - HWK EE TOOTH , d-TOOTH 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper In fallen block from Upper Bed I. Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper dentitions: Partial permanent and deciduous dentition. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 40 3755 78 - FLK S d-M 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Middle or Lower? Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper M2, broken. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sp. No
OH 41 3756 37 - HWK EE L-M 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper M1 or M2. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 42 3757 37 - HWK EE P-f 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Broken crown of upper premolar. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 5 3713 5 - FLK 283 - 45 CRA , MAX , M , I , C , P 17 July - 6 August 1959 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Middle (Zinj.) 22 feet below the upper limit of Bed I (Tuff IF), approximately 20 feet above basalt. Leakey, L. S. B. 1959. A new fossil skull from Olduvai. Nature 184:491-493. Holotype. Almost complete cranium. Adolescent. One reconstructed fossil found at House of Culture, Dar es Salaam, Tanzania. Specimen previously housed at National Museum of Kenya. Additional pieces from NMK unpublished. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Zinjanthropus boisei Holotype. This specimen was reclassified as Paranthropus boisei by Robinson (1960). No
OH 44 3759 5 - FLK R-M 1970 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Zinjanthropus floor Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Specimen is a crown of upper right M1. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 46 3761 6 - FLK NN TOOTH 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Broken crown of molar or premolar. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 1 3717 13 - RK 278 - 9 SKEL 1913 116 - Bayerische Staatssammlung für Paläontologie und Geologie (BSPG), Munich, Germany Ndutu Intrusive burial Hans-Reck. 1913, Oldoway-Expedition Monograph; LSB Leakey 18 March 1933 Partial skeleton. Intrusive burial post-dating primary Olduvai sequence. 144 - H. Reck 273 - Reck expedition Unknown Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sapiens No