Search the database

IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

Search Results

255 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
M. 21462 188 18 - VEK CRA, L/R-HOR 1931 106 - Natural History Museum UK (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The specimen is pictured in plate 85. It is pictured in plate 5 of Gentry (1966). 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Gazella This specimen was reclassified by Gentry and Gentry (1978) as Antidorcas recki. No
OVPP-Gazella 1 189 L/R-HOR 106 - Natural History Museum UK (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Gazella No
M. 14563 206 18 - VEK R-FRO, R-HOR 1931 106 - Natural History Museum UK (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 273 - Reck expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Neotragini Phenacotragus recki This specimen was reclassified as Gazella wellsi by Gentry (1966) and later to Antidorcas recki by Gentry and Gentry (1978). No
OVPP-Phenacotragus 1 207 11 - HWK UK 1935 106 - Natural History Museum UK (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The author did not state what the element is or the museum number. It was found at the base of Bed II. 130 - L.S.B. Leakey 273 - Reck expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Neotragini Phenacotragus recki No
M. 21429 211 18 - VEK CRA, L/R-HOR-f 1932 106 - Natural History Museum UK (?) II? Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The specimen was badly crushed prior to fossilization. It is pictured in plate 91. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini This specimen was reclassified as Parmularius cf. angusticornis by Gentry and Gentry (1978). Gentry and Gentry (1978) say this specimen's horn cores have attributes of P. antiquus as well. No
M. 21463 249 5 - FLK CRA, HOR 1935 106 - Natural History Museum UK (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is not described in the text but is shown in plate 84. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Gazella sp. This specimen was reclassified as Antidorcas recki by Gentry and Gentry (1978). No
M. 17090 253 10 - BK L-MAN-f, M 106 - Natural History Museum UK (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This holotype was described in a previous monograph (Leakey, 1958). 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Mesochoerus olduvaiensis Holotype No
M. 21484 868 51 - HEK L-HOR-f 1935 106 - Natural History Museum UK (?) II Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. 130 - L.S.B. Leakey No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros olduvensis This specimen was reclassified by Gentry and Gentry (1978) as Megalotragus kattwinkeli. No
M. 14683 10383 Two upper tusks, probably of the same individual. 1934 106 - Natural History Museum UK (?) I lower part, Middle Pleistocene Kamasian Pluvial LSB Leakey. 1958. Fossil Mammals of Africa: Some East African Pleistocene Suidae. British Museum. No. 14. British Museum Palaeontology Department 130 - L.S.B. Leakey 1942 Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Potamochoerus majus Paratypes. Originally classified as Koiropotamus majus (Hopwood, 1934). Reclassified as Potamochoerus majus (Leakey, 1942). Yes
M. 17071 10384 The greater part of a mandible. 1953 106 - Natural History Museum UK (?) II Chellean I Living Site LSB Leakey. 1958. Fossil Mammals of Africa: Some East African Pleistocene Suidae. British Museum. No. 14. marked Old. B.K.II, 1953/454; Brit. Mus. Palaeont. Dept. 130 - L.S.B. Leakey 1958 Yes Mammalia Artiodactyla Suiformes Suoidea Suidae Potamochoerus majus Yes
M. 17074 10385 A fragment of mandible with a well preserved lower third molar 1935 106 - Natural History Museum UK (?) I top of Bed I F.L.K. LSB Leakey. 1958. Fossil Mammals of Africa: Some East African Pleistocene Suidae. British Museum. No. 14. Brit. Mus. Palaeont. Dept. 130 - L.S.B. Leakey 1958 Unknown Mammalia Artiodactyla Suina Suoidea Suidae Potamochoerus majus Yes
M. 17073 10386 An isolated lower canine tooth. 1953 106 - Natural History Museum UK (?) II B.K.II LSB Leakey. 1958. Fossil Mammals of Africa: Some East African Pleistocene Suidae. British Museum. No. 14. Brit. Mus. Palaeont. Dept. Marked Old. B.K.II, 1953/443 130 - L.S.B. Leakey 1958 Unknown Mammalia Artiodactyla Suina Suoidea Suoidea Suidae Potamochoerus majus Yes
M. 17112 10387 An upper third molar in a fragment of maxilla. 106 - Natural History Museum UK (?) II F.L.K. Brit. Mus. Palaeont. Dept. 130 - L.S.B. Leakey 1958 Unknown Mammalia Artiodactyla Suina Suoidea Suidae Suidae Potamochoerus majus Yes
M. 17090 10388 A broken fragment of the left side of a mandible with damaged first molar, but with second and third molars intact. 106 - Natural History Museum UK (?) II Kamasian Pluvial LSB Leakey. 1958. Fossil Mammals of Africa: Some East African Pleistocene Suidae. British Museum. No. 14. Holotype. Formerly Coryndon Museum No. F. 3666. 130 - L.S.B. Leakey 1942 Unknown Mammalia Artiodactyla Suina Suoidea Suidae Suinae Suini Mesochoerus olduvaiensis Yes
263/1959 42 6 - FLK NN L/R-MAN, L/R-P, L/R-M, L/R-I, L/R-C 1959 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Hyaenidae Hyaeninae Crocuta ultra No
FLK I 506/1960 46 5 - FLK HUM, RAD, ULNA 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. All three bones are presumed by author to be from a single individual. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Hyaenidae Hyaeninae Crocuta ultra No
FLK N I 6140/1960 47 7 - FLK North HUM-f 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper 4 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Only the distal end of the humerus was recovered. It is severely crushed. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Carnivora Feliformia Hyaenidae Hyaeninae Crocuta ultra No
FLK N I 6307/1960 48 7 - FLK North ULNA-f 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper 4 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Only the proximal end of the ulna, much crushed, was recovered. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Hyaenidae Hyaeninae Crocuta ultra No
1510/1957 49 MAN-f, P, M 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Hyaenidae No
358/1960 50 SKULL, MAN, L/R-P, L/R-M 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Specimen is crushed. No locality is given. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Caniformia Cynoidea Canidae Canis mesomelas latirostris No
FLK N I 6252/1960 51 7 - FLK North SKULL, P, M 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Specimen is crushed. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Caniformia Cynoidea Canidae Canis mesomelas latirostris No
6237/1960 53 7 - FLK North L/R-RAM-f, L/R-P, L/R-M 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Caniformia Cynoidea Canidae Canis mesomelas latirostris No
24/1959 56 8 - MK M 1959 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I lower Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Carnivora Caniformia Arctoidea Mustelidae Lutrinae Lutra sp. No
1273/1957 57 MAN, L/R-P, L-C, L/R-M 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The author alludes to a skull and mandible that had not yet been described at the time of publication in 1965. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Felidae Pantherinae Panthera indet. No
301/1955 58 C 1955 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. No locality is given by author. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Felidae Pantherinae Panthera indet. No
302/1955 59 C-f 1955 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The canine is damaged. Author does not give information on the locality. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Felidae Pantherinae Panthera indet. No
1033/1957 60 P-f 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. No locality information is provided. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Felidae Pantherinae Panthera indet. No
791/1957 61 C-f 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This is only a damaged root of the canine. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Felidae Machairodontinae The author suggests it is like a saber-toothed cat but is "far too large to belong to the small species...from Bed 1." This specimen could be one of the specimens listed in Leakey (1971) from DK, FLK NN, FLK N II, or SHK. No
FLK NN I 177/1960 107 6 - FLK NN CRA, MAX, P, M, C 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle 3 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 28. This skull is "very crushed."Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Potamochoerus intermedius Holotype No
FLK N I 637 114 5 - FLK HOR 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle upper layers in the middle level. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Giraffoidea Giraffidae Palaeotraginae Okapia stillei Gentry and Gentry (1978) later identified this specimen as Parmularius altidens. No
F. 3665 P.P.T. 4 124 16 - LK CRA, L-HOR-f, R-HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Face and maxillae are missing. The holotype is pictured in plates 43 and 44. It was found on the surface between RK and LK. This eland is poorly represented at Olduvai. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Taurotragini Taurotragus arkelli Holotype No
OVPP-Gorgon 2 131 10 - BK HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Gorgon olduvaiensis No
P.P.R. 1 132 10 - BK FRLT, L/R-HOR 1953 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Reduncinae Reduncini Kobus idet. This specimen was reclassified by Gentry and Gentry (1978) as Kobus kob. No
579/1957 133 19 - SHK FRO-f, PAR-f, R-HOR 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Reduncinae Reduncini Kobus sp. No
OVPP-Hippotragus 1 136 HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Several specimens are present in the Nairobi Museum's collections. Three are pictured in plate 55. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus niro No
53/282 137 FRO, L/R-HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus sp. No
F. 3662 068/5812 P.P.T.2 138 CRA-f, L-HOR, R-HOR-f 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 56 and 58. The adhering matrix suggests this specimen must come from above the Lemuta member. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas Holotype No
1957.56 P.P.T.3 139 18 - VEK CRA-f, L/R-HOR 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper Upper Villafranchian Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 59 and 60. It is from a female. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas Paratype No
F. 992 141 HOR-f 1941 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas No
281/1953 142 19 - SHK HOR 1953 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas No