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IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

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2823 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
M. 21446 179 L-HOR-f, CRA-f 1935 106 - Natural History Museum UK (?) I Upper Villafranchian Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 80. Additional specimens have been found at FLK N I. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Beatragus antiquus Paratype No
M. 14950 177 22 - SC FRLT, L-HOR, R-HOR-f 106 - Natural History Museum UK (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 79 of Leakey (1965). 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Antidorcas kattwinkelli Holotype - This specimen was reclassified by Gentry and Gentry (1978) as Megalotragus kattwinkeli. No
F. 3013 P.P.F. 6 176 L/R-HOR-f, CRA-f, FRLT 1941 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 78. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Antidorcas kattwinkeli This specimen was reclassified by Gentry and Gentry (1978) as Megalotragus kattwinkeli. No
F. 3013 P.P.F. 6 4306 L/R-HOR-f, CRA-f, FRLT 1941 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 78. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Antidorcas kattwinkeli This specimen was reclassified by Gentry and Gentry (1978) as Megalotragus kattwinkeli. No
F. 3001 P.P.F. 5 175 CRA-f, HOR-f 1941 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 77. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 137 - A. Gentry 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius angusticornis This specimen was reclassified by Gentry and Gentry (1978) as Parmularius sp. indet. No
1315/1960 167 7 - FLK North L/R-HOR-f 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper All levels Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 73. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius altidens No
Ba. 126/1960 166 5 - FLK FRLT, R-HOR, L-HOR-f 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle Zinjanthropus (22 feet below the upper limit of bed I) layer and upper layers. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 72. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius altidens No
M. 14514 165 CRA-f, HOR 1931 106 - Natural History Museum UK (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 71. 293 - A.T. Hopwood 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius altidens No
G. 390 151 5 - FLK L-HOR, L-FRLT, FRO 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 62. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Oryx indet. This specimen was reclassified by Gentry and Gentry (1978) as Oryx gazella. No
F. 3000 219 19 - SHK R-FRLT, R-HOR-f 1941 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 40, figure 2. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Caprinae Caprini This specimen was reclassified as an unidentified Alcelaphini by Gentry and Gentry (1978). No
S. 217 1093 14 - HWK E FRLT, L-HOR, R/HOR-f 1971 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II rootlet clay Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 34, figure 1. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 137 - A. Gentry 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Beatragus antiquus No
S. 208 1099 CRA, PAL-f, L-M 1970 104 - Unknown Lemuta Tuff Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 32, figure 1. 137 - A. Gentry Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Indet. This specimen has some resemblance to Rabaticeras arambourgi. It is a different species than specimen 1963.068/5976. No
JK II A. 1129 1079 46 - JK 2 FRLT, R-HOR-f 104 - Unknown III Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 31, figure 1. 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Rabaticeras arambourgi No
068/6514 3711 34 - MLK CRA, MAX-f, R/L-P, R/L-M 104 - Unknown II middle Leakey, M. G., and Leakey, R. E. F. 1973. Pleistocene Colobinae from East Africa. Leakey, L. S. B., Savage, R. J. G., Coryndon, S. C., eds. In: Fossil Vertebrates of Africa,vol. 3. London: Academic Pr This specimen is pictured in plate 3. 154 - Leakey and Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Colobinae Colobini Cercopithecoides kimeui Holotype No
01. R. L5-58 3128 7 - FLK North L-MAX-f, M 104 - Unknown I 5 Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 3, figures G and H. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Praomys minor No
01. R. L2-2 3127 L-MAX-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 3, figures A and B. This species is found at FLK NI, layers 1, 2, 3, 4, 5, and 6; FLK NNI, layers 1, 2, and 3; FLK I, Zinjanthropus layer. It is unclear where individual specimens come from, however. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Mus petteri Holotype No
01. R. K-314 3130 5 - FLK R-MAX-f, M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 3, figure D. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Zelotomys leakeyi Holotype No
Pel 23 5758 MAX-f, PMAX-f 1957 122 - National Museum and House of Culture, Dar es Salaam, Tanzania Gentry, A. W. 1967. Pelorovis oldowayensis Reck, an extinct bovid from East Africa: British Museum. pp. 245-299 This specimen is pictured in plate 3, figure 5. 137 - A. Gentry 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Pelorovis oldowayensis No
Pel 23 1691 MAX-f, PMAX-f 1957 122 - National Museum and House of Culture, Dar es Salaam, Tanzania Gentry, A. W. 1967. Pelorovis oldowayensis Reck, an extinct bovid from East Africa: British Museum. pp. 245-299 This specimen is pictured in plate 3, figure 5. 137 - A. Gentry 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Pelorovis oldowayensis No
1957 SHK II 232 1692 19 - SHK MAX-f 1957 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Gentry, A. W. 1967. Pelorovis oldowayensis Reck, an extinct bovid from East Africa: British Museum. pp. 245-299 This specimen is pictured in plate 3, figure 4. 137 - A. Gentry 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Pelorovis oldowayensis No
1952 BK II 119 1699 10 - BK L-MAN-f, L-M, L-P 1952 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Gentry, A. W. 1967. Pelorovis oldowayensis Reck, an extinct bovid from East Africa: British Museum. pp. 245-299 This specimen is pictured in plate 3, figure 3. 137 - A. Gentry 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Pelorovis oldowayensis No
1952 BK II 117 1697 10 - BK L-MAN-f, L-M, L-P 1952 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Gentry, A. W. 1967. Pelorovis oldowayensis Reck, an extinct bovid from East Africa: British Museum. pp. 245-299 This specimen is pictured in plate 3, figure 2. 137 - A. Gentry 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Pelorovis oldowayensis No
HWK EE II 1965.068/5975 1018 37 - HWK EE CRA-f, R/L-HOR 1965 104 - Unknown II middle found 3.7 meters above the Sandy Conglomerate Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 26, figures 1, 2. 131 - A. Remane Yes "pyroxene-rich volcanic sandstone" Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius sp. No
BK II 1963.2499 991 10 - BK FRLT-f, R-HOR 1963 104 - Unknown II Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 25, figure 3. 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius angusticornis No
1960.067/4951 1011 6 - FLK NN CRA, HOR-F 1960 104 - Unknown I 10 feet above the base of Bed 2 Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 25, figure 2. The horn core fragments are registered as: 1960.067/4949 067/4946 067/4948 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius angusticornis This specimen represents an intermediate stage of P. angusticornis and P. altidens. No
HWK I 1962.068/6650 936 11 - HWK CRA-f, HOR 1962 104 - Unknown I Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 20, figure 1. 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius altidens No
01. R. K-199 3124 5 - FLK R-MAX-f, M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figures A and B. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus Holotype No
01. R. K-533 3125 5 - FLK R-MAX-f, M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figure E. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Aethomys lavocati Holotype No
01. R. M5-223 3353 L-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figure D. This species is found at FLK NNI layer 2 and FLK NI layers 1, 2, 4, and 5. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Pelomys dietrichi No
01. R. M5-222 3126 7 - FLK North R-MAX-f, M 104 - Unknown I 5 Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figure C. 153 - J. Jaeger Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Pelomys dietrichi Holotype No
01. R. K-200 3331 R-MAN-f, M 104 - Unknown I Jaeger, J.J. 1976. Les rongeurs (Mammalia, Rodentia) du Pleistocene inferieur d’Olduvai Bed I (Tanzanie) 1ere Partie: Les Murides. Fossil vertebrates of Africa 4:58-120. This specimen is pictured in plate 2, figure B. This species is found at FLK I; FLK NNI layers 2 and 3; FLK NI layer 4. It is unclear where individual specimens were recovered. 153 - J. Jaeger Unknown Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Murinae Arvicanthis primaevus No
OVPP-Elephas 5 251 M-f 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 16. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 264 - Leakey family expedition Unknown Mammalia Proboscidea Tethytheria Behemota Elephantoidea Elephantidae Elephas africanavus No
SHK II 1957.946 1293 19 - SHK HOR-f 1957 104 - Unknown II Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 16, figure 2 (right). 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Connochaetes sp. No
MNK II 2716 902 27 - MNK L-HOR-f 104 - Unknown II Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 16, figure 1. 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Connochaetes taurinus olduvaiensis The authors suggest this specimen may be from C. taurinus and if so, would be the earliest of this subspecies at Olduvai. No
M. 14681 1313 L-MAX-f, L-M, L-P 106 - Natural History Museum UK (?) II Hopwood, A. 1934. New fossil mammals from Olduvai, Tanganyika territory. Annals of the Magazine of Natural History. 10: 546-550. This specimen is pictured in plate 14b and 14c (Jolly, 1972). 293 - A.T. Hopwood Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Cynocephalinae Simopithecus leakeyi This specimen was reclassified as aff. Theropithecus by Jolly (1972). No
GTC IV 1962.068/6664 874 35 - GTC R-HOR, L-HOR-f, FRLT 1962 104 - Unknown IV Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 13, figure 1. Note, the teeth shown in the top of the image are from a different specimen. 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Megalotragus kattwinkelli No
MNK II 3258 867 27 - MNK L-HOR, CRA-f, L-FRO 104 - Unknown II Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 12. 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Megalotragus kattwinkelli No
HWK East II 131 1092 14 - HWK E R-HOR-f 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II 1 Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 12, figure 1. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 137 - A. Gentry 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Beatragus antiquus No
BK II 1963.2226 841 10 - BK L-MAX-f, L-M 1963 104 - Unknown II Gentry, A. W., and Gentry, A. 1978. Fossil Bovidae (Mammalia) of Olduvai Gorge, Tanzania. Part I. Bulletin of the British Museum (Natural History): Geology 29:289-446. This specimen is pictured in plate 11, figure 4 137 - A. Gentry Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas No
S. 196 3116 24 - DK CRA-f, P, M, C 1969 104 - Unknown I Leakey, M., and Leakey, R. 1976. Further Cercopithecinae (Mammalia, Primates) from the Plio/Pleistocene of East Africa. Fossil vertebrates of Africa 4:121-146. This specimen is pictured in plate 1. 142 - M. D. Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Cercopithecinae Papionini Papio ap. No