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IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

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1427 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
OH 11 3726 24 - DK 285 - 13 PAL , L-MAX-f , TOOTH-f 1962 122 - National Museum and House of Culture, Dar es Salaam, Tanzania Ndutu Lower Found in stream bed at bottom of gully near DK excavations. See curatorial notes for details. Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left half of palate and maxillary arch, crowns of teeth not preserved. "It is thus impossible to assign to any stratigraphic level, although the condition of the bone, which bears a pinkish tinge, is similar to that seen in fossil bones from a conglomerate at the head of the DK gully belonging to the lower Ndutu Beds" (Leakey MD, 1971:230). Specimen previously housed at National Museum of Kenya. 154 - Leakey and Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sp. No
OH 12 3727 18 - VEK 286 - 86 OCC , L/R-PAR , L/R-TEM-f , CRA-f , L-PAL , L-MAX , TOOTH-f 1962 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV? a Surface. Found on eroded slope of Bed III. L-TEM-f from just below marker Tuff IVa. Leakey, L., and Leakey, M. 1964. Recent discoveries of fossil hominids in tanganyika: at Olduvai and near Lake Natron. Nature 202:5-7. Crowns of teeth not preserved. Bed IVA determined on basis of matrix adhering to L-TEM-f and PAL. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No L-TEM-f & PAL found with matrix adhering. Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 13 3728 27 - MNK 287 - 88 "Skull Site" MAN , MAX , CRA-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle IIB 5 feet above chert horizon Paratpye. "This consists of a sub-adult skull, probably female, with the third molars coming into wear. It is represented by the greater part of the mandible, including the complete dentition, although the crowns of the incisors are damaged and the left ascending ram us is not preserved; the greater part of the maxillae, represented by two fragments both lacking the anterior portions; the greater part of the occipital; the right parietal and temporal; parts of the left parietal and temporal; together with a few frontal fragments and approximately fifty additional pieces which it has not been possible to fit together" (Leakey MD, 1971:231). Specimen previously housed at National Museum of Kenya. 5 pieces found at House of Culture, Dar es Salaam, Tanzania. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype No
OH 14 3729 27 - MNK 287 - 88 "Skull Site" CRA-f , FRO-f , R-PAR-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle B Surface. Inferred as same horizon as OH 13 (Leakey MD, 1971). Leakey, L.S.B., Tobias, P.V. & Napier, J.R. 1964. A new species of the genus Homo from Olduvai Gorge. Nature 202(4927): 7-9. Juvenile (sutures unfused). Thin skull. Specimen previously housed at National Museum of Kenya. 11 pieces found at House of Culture, Dar es Salaam, Tanzania. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Juvenile, parietal is thin. No
OH 15 3730 27 - MNK 287 - 88 "Skull Site" C , M 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle Above level of OH 13. Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. One canine, two molars. Canine is worn, molars are upper molars. "The teeth are exceedingly robust and show an unusual degree of wear. They were not found in close association; one molar and the canine lay approximately I ft. "Qelow the top of the tuff in which they occurred, whilst the second molar was 4 ft. distant and 1 ft. deeper in the deposit" (Leakey MD, 1971:232). Specimen previously housed at National Museum of Kenya. 7 pieces found at House of Culture, Dar es Salaam, Tanzania. 154 - Leakey and Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo No
OH 16 3731 72 - FLK Maiko Gully 283 - 45 CRA-f , TOOTH , MAN , 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II base Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Upper and lower dentition, calotte, and > 1500 unidentified fragments due to cattle trampling. Majority of fragments found on surface of Tuff IF, some teeth found in situ, 2 feet above Tuff IF. Specimen previously housed at National Museum of Kenya. 154 - Leakey and Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Leakey LSB, 1964 refers to this specimen as Homo erectus. Leakey, Tobias, and Napier provisionally assign it to Homo habilis (referred material, not paratype) (1964), and Leakey MD, 1971 refers to it as cf. Australopithecus. Yes
OH 17 3732 72 - FLK Maiko Gully d-M 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Surface Stream bed of Maiko Gully Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 19 3734 29 - FC W 288 - 89 M-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Middle 15 feet above chert horizon Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Half of a very worn molar. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 20 3735 11 - HWK 289 - 44 L-FEM-f 1959 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II or I Lowermost if Bed II, upper if Bed I Surface Day, MH. 1969. Femoral Fragment of a Robust Australopithecine from Olduvai Gorge, Tanzania. Nature. 22: 230-233. Proximal end of large left femur missing head, greater and lesser trochanters damaged. Either Lower Bed II or Upper Bed I (Leakey MD, 1971:233). Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus cf. boisei No
OH 21 3736 7 - FLK North L-M 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper Surface Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Upper left M1, small and unworn. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 22 3737 73 - VEK/MNK R-MAN-f , R-P , R-M , R-I-f , R-C-f 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania III/IV Surface, indeterminate gravel Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Right half of mandible with roots of I1-C and complete P3-M2. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo cf. erectus No
OH 23 3738 5 - FLK L-MAN-f , L-P , L-M , 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV/Masek Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left mandible fragment with P4-M2. Specimen previously housed at National Museum of Kenya. Bed IV in Leakey (1971), referred to as having derived from Masek Beds in other publications (e.g., Millard, 2009). 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 24 3739 74 - DK E CRA , M , P 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Lower Surface Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Cranium is crushed. Teeth not specified. From same horizon as OH 4. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 25 3740 268 - 54 L-PAR-f 1968 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV a Surface Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left parietal fragment ca. asterion. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Juvenile. No
OH 26 3741 55 - FLK W R-M 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I/II See curatorial notes. Upper Surface, .6 m below Tuff If at top of Bed I. Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Large unworn right L. M3. "Direct evidence as to the original position of the tooth was ... not obtained but, on the basis of the adhering matrix, which is indistinguishable from the limestone nodules overlying Tuff IF, it can be assumed to have come from the base of Bed II..." (Tobias, 1991:848). Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Found embedded in small nodule of calcareous concretion. Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 27 3742 37 - HWK EE R-M 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 meters below Tuff IF Surface. Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Lower M3, unworn. "It is not possible to indicate more than broadly the original position of this tooth, since the clays above and below Tuff IF, as well as the Sandy Conglomerate, are all fossiliferous in this areaand yield mammalian teeth in a similar state of preservation to the hominid molar...the tooth was derived from either Upper Bed I or Lower Bed II" (Tobias, 1991:849). Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 28 3743 38 - WK 290 - 36 L-PVG-f , L-FEM-f 1970 122 - National Museum and House of Culture, Dar es Salaam, Tanzania IV a Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Left femoral shaft and left innominate. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 29 3744 76 - JK W M-f 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania III/IVa Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 30 3745 72 - FLK Maiko Gully 283 - 45 TOOTH , d-TOOTH , CRA-f , 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Surface "Parts deciduous and germ dentitions" (Leakey MD, 1971:234). Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 31 3746 14 - HWK E M-f 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper Surface 1.5 m below Tuff IF Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo No
OH 32 3747 27 - MNK M-f 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Middle Surface. Broken. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo No
OH 33 3748 6 - FLK NN CRA-f 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Surface Tobias, PV. 1991. The Skulls, endocasts and teeth of Homo habilis. Cambridge: University Press. Thin vault fragments. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo Very thin skull No
OH 34 3749 76 - JK W FEM , TIB-f 1962 122 - National Museum and House of Culture, Dar es Salaam, Tanzania III/ IV for Bed IV level = a Leakey MD. (1971) Olduvai Gorge, Vol. 3: excavations in Beds I & II 1960-1963. New York, NY: Cambridge University Press. Femur described as "slender" (Leakey MD, 1971) and shaft of tibia. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sp. Slender bone No
OH 35 3750 6 - FLK NN TIB , FIB 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 22 Davis, P. R. 1964. Hominid fossils from Bed I, Olduvai Gorge, Tanganyika. A tibia and fibula. Nature. 201, 967. Almost complete. Attributed to same individual as skull fragments and teeth of OH 6 but appear to be more mature (Tobias, 1991). Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 36 3751 22 - SC ULNA 1970 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Upper Tuff IID Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Almost complete. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 37 3752 5 - FLK L-MAN , M , M 1971 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Middle Surface, probably from sandy conglomerate in Middle Bed II. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Left half mandible with M1-M2. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 38 3753 22 - SC R-M , I , I 1971 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Upper >IID Fluvo-lacustrine Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Molar is right M2. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 39 3754 37 - HWK EE TOOTH , d-TOOTH 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper In fallen block from Upper Bed I. Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper dentitions: Partial permanent and deciduous dentition. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 40 3755 78 - FLK S d-M 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Middle or Lower? Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper M2, broken. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo sp. No
OH 41 3756 37 - HWK EE L-M 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper M1 or M2. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 42 3757 37 - HWK EE P-f 1972 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Upper Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Broken crown of upper premolar. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 43 3758 6 - FLK NN L-MTT, L-MTT 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 44 3759 5 - FLK R-M 1970 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Zinjanthropus floor Surface. Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Specimen is a crown of upper right M1. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 45 3760 6 - FLK NN L-M 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Upper M1 germ. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 46 3761 6 - FLK NN TOOTH 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 Leakey, M.D. 1978. Olduvai fossil hominids:their stratigraphic positions and associations. In: Jolly, CJ, editor. Early hominids of Africa. London:Duckworth. P. 3-16. Broken crown of molar or premolar. Specimen previously housed at National Museum of Kenya. 142 - M. D. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Australopithecus boisei No
OH 48 3763 6 - FLK NN CLA 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 -- Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 49 3764 6 - FLK NN RAD-f 1960 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 -- Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No
OH 50 3765 6 - FLK NN RIB-f 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I 3 -- Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae No
OH 51 3766 35 - GTC L-MAND-f, L-P, L-M 1974 122 - National Museum and House of Culture, Dar es Salaam, Tanzania III Surface. -- Mandible fragment with P4 and M1. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo erectus No
OH 52 3767 24 - DK L-CRA-f 1969 122 - National Museum and House of Culture, Dar es Salaam, Tanzania I Lower Surface. -- Incomplete left temporal. Specimen previously housed at National Museum of Kenya. 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis No