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IMPORTANT NOTE: We have maintained the originally published taxonomic designation for each specimen despite the fact that many of these specimens have been reclassified since.  We chose to follow this protocol because of the difficulty in keeping up to date with all of the taxonomic groups, and our hesitancy to make a judgement call on taxonomic debates.  As such, please search the database with this in mind, perhaps using higher level taxonomic classifications for your query.

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255 specimens found
Image Spec # CODI Database ID Arch. Locality Geo. Locality Elements Preserved Excavation Date Repository Bed Level Stratigraphic Horizon Bibliographic Information Curatorial Notes ID By ID Date Research Team In Situ Sediment or Matrix Adhering Class Order Suborder Infraorder Superfamily Family Subfamily Tribe Genus Species Subspecies Taxonomic Notes Taxonomic Problems
M. 21470 117 12 - CK R-HOR-f 1935 106 - Natural History Museum UK (?) Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros sp. This specimen was reclassified as Tragelaphus stepsiceros by Gentry and Gentry (1978). No
M. 21469 120 12 - CK R-HOR-f 1935 106 - Natural History Museum UK (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. No information is given about elements preserved. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros grandis Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros grandis. No
M. 21461 118 13 - RK CRA, R/L-HOR, R-I, R-P, R-M 1931 106 - Natural History Museum UK (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 38 and 39. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros grandis Holotype. Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros grandis. No
OVPP-Tragelaphus 1 121 14 - HWK E CRA, HOR 1959 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II lower 2 Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 40, 41. The cranium and horn cones were found near the base of Bed 2. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros maryanus Holotype - Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros maryanus. No
068/5813 123 58 - KK CRA 1959 104 - Unknown II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The author indicates that specimens are common in Bed I, lower Bed II, and early in middle Bed II (Gentry and Gentry, 1978, a). The specimen # is provided by Gentry and Gentry (1978). The cranium and horn cones were found near the base of Bed 2. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Strepsicerotini Strepsiceros maryanus Gentry and Gentry (1978) reclassified this specimen as Tragelaphus strepsiceros maryanus. No
F. 3665 P.P.T. 4 124 16 - LK CRA, L-HOR-f, R-HOR 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Face and maxillae are missing. The holotype is pictured in plates 43 and 44. It was found on the surface between RK and LK. This eland is poorly represented at Olduvai. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Taurotragini Taurotragus arkelli Holotype No
M. 14947 125 17 - PLK FRLT-f, L/R-HOR-f 106 - Natural History Museum UK (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Specimen is pictured in plate 46. According to the author, it is "in bad shape." Gentry and Gentry suggest this specimen probably came from the top of Bed 2 at PLK. 130 - L.S.B. Leakey 1936 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Bularchus arok Holotype - Gentry and Gentry (1978) reclassidfied this specimen as Pelorovis oldowayensis. No
M. 21451 128 18 - VEK CRA-f, R-HOR-f 1935 106 - Natural History Museum UK (?) III, IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This holotype was found at the junction of Beds 3 and 4. It is pictured in plates 49 and 50. [This species is documented at SHK II, FLK N I, and BK II (Leakey, 1971).] 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Bovinae Bovini Gorgon olduvaiensis Holotype - Gentry and Gentry (1978) reclassified this specimen as Connochaetes taurinus olduvaiensis. No
1957.56 P.P.T.3 139 18 - VEK CRA-f, L/R-HOR 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I upper Upper Villafranchian Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 59 and 60. It is from a female. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas Paratype No
M. 21462 188 18 - VEK CRA, L/R-HOR 1931 106 - Natural History Museum UK (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The specimen is pictured in plate 85. It is pictured in plate 5 of Gentry (1966). 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Gazella This specimen was reclassified by Gentry and Gentry (1978) as Antidorcas recki. No
M. 14563 206 18 - VEK R-FRO, R-HOR 1931 106 - Natural History Museum UK (?) IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 273 - Reck expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Neotragini Phenacotragus recki This specimen was reclassified as Gazella wellsi by Gentry (1966) and later to Antidorcas recki by Gentry and Gentry (1978). No
M. 21429 211 18 - VEK CRA, L/R-HOR-f 1932 106 - Natural History Museum UK (?) II? Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. The specimen was badly crushed prior to fossilization. It is pictured in plate 91. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini This specimen was reclassified as Parmularius cf. angusticornis by Gentry and Gentry (1978). Gentry and Gentry (1978) say this specimen's horn cores have attributes of P. antiquus as well. No
579/1957 133 19 - SHK FRO-f, PAR-f, R-HOR 1957 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Reduncinae Reduncini Kobus sp. No
281/1953 142 19 - SHK HOR 1953 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Hippotragus gigas No
F. 3000 219 19 - SHK R-FRLT, R-HOR-f 1941 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 40, figure 2. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Caprinae Caprini This specimen was reclassified as an unidentified Alcelaphini by Gentry and Gentry (1978). No
SHK II 1953.234 220 19 - SHK HOR-f 1953 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Caprinae Caprini This specimen was reclassified as an unidentified Alcelaphini by Gentry and Gentry (1978). No
OVPP-Clarias 2 227 19 - SHK UK 104 - Unknown II Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Actinopterygii Siluriformes Clariidae Clarias sp. No
M. 21430 174 21 - HWK Castle SKULL, R-HOR-f 1932 106 - Natural History Museum UK (?) IV Base of Bed IV. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plates 75 and 76. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius rugosus Holotype No
M. 14950 177 22 - SC FRLT, L-HOR, R-HOR-f 106 - Natural History Museum UK (?) II upper Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 79 of Leakey (1965). 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Antidorcas kattwinkelli Holotype - This specimen was reclassified by Gentry and Gentry (1978) as Megalotragus kattwinkeli. No
OLD 580 3695 22 - SC MAN-f, M 1957 104 - Unknown II Leakey, L., and Whitworth, T. 1958. Notes on the genus Simopithecus, with a description of a new species from Olduvai. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Primates Anthropoidea Haplorhini Cercopithecoidea Cercopithecidae Cynocephalinae Simopithecus jonathani No
M. 21447 181 23 - TK CRA, L/R-MAX-f, L-HOR, L/R-P, L/R-M, L/R-I 1931 106 - Natural History Museum UK (?) III, IV 4 feet below Masek Beds Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This neotype specimen is pictured in plates 81 and 82. The original holotype of M. kattwinkelli (VI-1099) was lost in WWII in Munich. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Xenocephalus kattwinkelli Neotype - This specimen was reclassified by Gentry and Gentry (1978) as Megalotragus kattwinkeli. No
M. 22360 187 24 - DK R-HOR 1935 106 - Natural History Museum UK (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen was originally assigned to aff. Gazella by Leakey (1965) and later assigned to the species G. wellsi (Gentry, 1966) and finally to Antidorcas recki by Gentry and Gentry (1978a). 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Antilopinae Antilopini Gazella This specimen was reclassified as Gazella wellsi by Gentry (1966) and later by Gentry and Gentry (1978) as Antidorcas recki. No
OVPP-Hippopotamus 2 282 24 - DK UK 104 - Unknown I lower All levels 130 - L.S.B. Leakey Unknown Mammalia Artiodactyla Suiformes Hippopotamoidea Hippopotamidae Hippopotaminae Hippopotamus gorgops No
OVPP-Tilapia 2 311 24 - DK UK 104 - Unknown I lower 130 - L.S.B. Leakey Unknown Actinopterygii Perciformes Cichlidae Pseudocrenilabrinae Tilapini Tilapia sp. No
OVPP-Serpentes 3 234 26 - FC VER 104 - Unknown IV Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Several snake specimens have been found. 130 - L.S.B. Leakey 264 - Leakey family expedition No Reptilia Serpentes No
OH 13 3728 27 - MNK 287 - 88 "Skull Site" MAN , MAX , CRA-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle IIB 5 feet above chert horizon Paratpye. "This consists of a sub-adult skull, probably female, with the third molars coming into wear. It is represented by the greater part of the mandible, including the complete dentition, although the crowns of the incisors are damaged and the left ascending ram us is not preserved; the greater part of the maxillae, represented by two fragments both lacking the anterior portions; the greater part of the occipital; the right parietal and temporal; parts of the left parietal and temporal; together with a few frontal fragments and approximately fifty additional pieces which it has not been possible to fit together" (Leakey MD, 1971:231). Specimen previously housed at National Museum of Kenya. 5 pieces found at House of Culture, Dar es Salaam, Tanzania. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Homininae Hominini Homo habilis Paratype No
OH 14 3729 27 - MNK 287 - 88 "Skull Site" CRA-f , FRO-f , R-PAR-f 1963 122 - National Museum and House of Culture, Dar es Salaam, Tanzania II Lower Middle B Surface. Inferred as same horizon as OH 13 (Leakey MD, 1971). Leakey, L.S.B., Tobias, P.V. & Napier, J.R. 1964. A new species of the genus Homo from Olduvai Gorge. Nature 202(4927): 7-9. Juvenile (sutures unfused). Thin skull. Specimen previously housed at National Museum of Kenya. 11 pieces found at House of Culture, Dar es Salaam, Tanzania. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Primates Anthropoidea Haplorhini Hominoidea Hominidae Juvenile, parietal is thin. No
FLK I/E-G 393 4 5 - FLK L-RAM, L-P, L-M 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Primates Strepsirrhini Lorisiformes Galagidae Galago senegalensis No
OVPP-Suncus 4 13 5 - FLK UK 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Only one specimen is found in the Zinjanthropus level (22). 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Insectivora Soricoidea Soricidae Crocidurinae Suncus orangiae No
OVPP-Gerbillus 1 27 5 - FLK UK 104 - Unknown I 5, 6 and perhaps above but not explicitly stated. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Elements preserved are not indicated by the author. 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Gerbilinae Gerbilini Gerbillus sp. No
OVPP-Tatera 1 25 5 - FLK 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Several specimens are present. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae Gerbillinae Taterillini Tatera sp. No
OVPP-Muridae 1 35 5 - FLK P, M, I, C, MAN, MAX 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Several unidentified murids 130 - L.S.B. Leakey 264 - Leakey family expedition No Mammalia Rodentia Myomorpha Myodonta Muroidea Muridae No
FLK I 506/1960 46 5 - FLK HUM, RAD, ULNA 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. All three bones are presumed by author to be from a single individual. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Carnivora Feliformia Hyaenidae Hyaeninae Crocuta ultra No
G. 356 102 5 - FLK MAN, L/R-P, L/R-M, L/R-C, L/R-I 1960 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 23. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Suini Promesochoerus mukiri Holotype No
OVPP-Tapinochoerus 1 110 5 - FLK R-MAN, R-P, R-M 1959 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 30. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Phacochoerini Tapinochoerus sp. No
OLD 159 111 5 - FLK M 1959 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 31. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Suiformes Suoidea Suidae Suinae Phacochoerini Tapinochoerus No
FLK N I 637 114 5 - FLK HOR 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle upper layers in the middle level. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Giraffoidea Giraffidae Palaeotraginae Okapia stillei Gentry and Gentry (1978) later identified this specimen as Parmularius altidens. No
G. 390 151 5 - FLK L-HOR, L-FRLT, FRO 104 - Unknown I middle (Zinjanthropus) 22 feet below the upper limit of bed I. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 62. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Hippotraginae Hippotragini Oryx indet. This specimen was reclassified by Gentry and Gentry (1978) as Oryx gazella. No
Ba. 126/1960 166 5 - FLK FRLT, R-HOR, L-HOR-f 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I middle Zinjanthropus (22 feet below the upper limit of bed I) layer and upper layers. Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 72. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Yes Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius altidens No
F. 206/1960 P.P.F. 7 168 5 - FLK HOR 1960 105 - National Museum and House of Culture, Dar es Salaam, Tanzania (?) I Leakey LSB. 1965. Olduvai Gorge 1951-1961, Volume 1: Fauna and Background. New York, NY: Cambridge University Press. This specimen is pictured in plate 74. Original information indicated that this specimen was located in Nairobi. This specimen is likely within the material that was repatriated to Dar in 2011, but this has yet to be confirmed. 130 - L.S.B. Leakey 264 - Leakey family expedition Unknown Mammalia Artiodactyla Ruminantia Bovoidea Bovidae Alcelaphinae Alcelaphini Parmularius altidens Unknown